Google is now better able to reward high-quality Web content that is relevant to a search query, and less dependent on traditional signals, such as keywords, according to a recent report from Searchmetrics.

The report was based on an analysis of the the top 30 search results for 10,000 relevant keywords and 300,000 websites appearing on in 2015. The aim of the study, which has been carried out every year since 2012, was to identify the key factors that high-ranking Web pages have in common.

Here are four key search engine optimization (SEO) lessons from the report.